Yes, if the rifle says it is chambered for both, it will take both.
The rounds are practically the same, but some barrels are more exact than others.
For example, the military-spec AR-15 is chambered for the 5.56, and though you might be able to stuff a .223 in there, it can cause problems.
Different but similar calibers usually come about as the result of military trials, where they try to fine tune the weapon, plus they want a caliber specific to NATO standards.
You'll find civilian-grade weapons are often more forgiving.




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